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Ikuo Kandori


FacultyDepartment of Agricultural Science / Graduate School of Agluculture
PositionAssociate Professor
Commentator Guide
Last Updated :2018/10/14

Education and Career


  •  - 1998 , Kyoto University, Graduate School, Division of Agriculture
  •  - 1991 , Kyoto University, Faculty of Agriculture

Research Activities

Research Areas

  • Basic biology, Ecology/Environment, Ecological Entomology
  • Boundary agriculture, Insect science, Applied Entomology

Research Interests

  • Pollination Ecology, Ecological Entomology

Published Papers

  • The choosing of sleeping position in the overnight aggregation by the solitary bees Amegilla florea urens in Iriomote Island of Japan., Yokoi, T., Idogawa, N., Kandori, I., Nikkeshi, A. and Watanabe, M., The Science of Nature, 104, 23 - 23, Mar. 2017 , Refereed
  • Comparison of foraging activity between mason bee Osmia orientalis (Hymenoptera: Megachilidae) and honybees for wild raspberry Rubus hirsutus (Rosales: Rosaceae)., Yokoi, T. and Kandori, I., Entomological News, 125(5), 363 - 367, Mar. 2016 , Refereed
  • Long frontal projections help Battus philenor (Lepidoptera: Papilionidae) larvae find host plants, Ikuo Kandori, Kazuko Tsuchihara, Taichi A. Suzuki, Tomoyuki Yokoi, Daniel R. Papaj, PLoS One, 10(7), e0131596 - e0131596, Jul. 2015 , Refereed
  • Strongly biased sex ratio in cuckoo wasp Chrysura hirsuta (Hymenoptera: Chrysididae), a parasitoid of the mason bee Osmia orientalis , Yokoi Tomoyuki; Hirooka Takuto; Terada Takeshi; Sugimoto, Shusaku; Tamaru Mayumi; Satoh Sachiko; Kandori Ikuo, Entomological Science, 15(2), 250 - 252, Apr. 2012 , Refereed
  • Reward and non-reward learning of flower colours in the butterfly Byasa alcinous (Lepidoptera: Papilionidae)., Kandori I, Yamaki T, Die Naturwissenschaften, 99(9), 705 - 713, Sep. 2012 , Refereed
  • Nesting Habits of a Pollinator: Osmia orientalis (Hymenoptera: Megachilidae), KANDORI Ikuo, TAMARU Mayumi, YOKOI Tomoyuki, Japanese Journal of Applied Entomology and Zoology, 54(2), 77 - 84, 2010 , Refereed
  • Interspecific and intersexual learning rate differences in four butterfly species., Ikuo Kandori, Takafumi Yamaki, Sei-ichi Okuyama, Noboru Sakamoto and Tomoyuki Yokoi, The Journal of Experimental Biology, 212(Pt 23), 3810 - 3816, Dec. 2009 , Refereed
  • An invasive dandelion unilaterally reduces the reproduction of a native congener through competition for pollination, Ikuo Kandori · Toshihiro Hirao · Satoshi Matsunaga · Tsutomu Kurosaki, Oecologia, 159(3), 559 - 569, Mar. 2009 , Refereed
  • Differences in growth and development of the parasitoids Dacnusa nipponica and D. sibirica (Hymenoptera: Braconidae) on the leaf-mining fly Liriomyza trifolii (Diptera: Agromyzidae) compared to various-sized leaf-mining Chromatomyia horticola hosts (Di・・・, Ikuo KANDORI, Masaki YAMADA, Tomoo HONDO and Tuyosi SUGIMOTO, Applied Entomology and Zoology, 43(4), 577 - 583, 2008 , Refereed
    Summary:Differences in growth and development of the parasitoids Dacnusa nipponica and D. sibirica (Hymenoptera: Braconidae) on the leaf-mining fly Liriomyza trifolii (Diptera: Agromyzidae) compared to various-sized leaf-mining Chromatomyia horticola hosts (Diptera: Agromyzidae)
  • Cold tolerance of the sweet potato weevil, Cylas formicarius (Fabricius) (Coleoptera: Brentidae), from the Southwestern Islands of Japan, Ikuo KANDORI, Toshio KIMURA, Hisaaki TSUMUKI and Tuyosi SUGIMOTO, Applied Entomology and Zoology, 41(2), 217 - 226, 2006 , Refereed
  • Diverse visitors with various pollinator importance and temporal change in the important pollinators of Geranium thunbergii (Geraniaceae), KANDORI Ikuo, Ecological Research, 17, 283 - 294, 2002 , Refereed
  • Effect of experience on foraging behavior towards artificial nectar guide in the cabbage butterfly, ┣DBPieris(/)-┫DB ┣DBrapae(/)-┫DB ┣DBcrucivora(/)-┫DB (Lepidoptera : Pieridae)., Applied Entomology and Zoology, 33(1), 35 - 42, 1998 , Refereed
  • The learning abilities of the white cabbage butterfly, ┣DBPieris(/)-┫DB ┣DBrapae(/)-┫DB, foraging for flowers., Researches on Population Ecology, 38(1), 111 - 117, 1996 , Refereed
  • Male mating behavior in ralation to spermatophore transfer in white cabbage butterfly., Researches on Population Ecology, 38(2), 225 - 230, 1996 , Refereed
  • Flower visitors and pests of mango in the hothouse, 38, 19 - 30, Mar. 2005
  • On elytral color dimorphism of sweet potato weevil, Cylas formicarius (Fabricius), in the Southwest islands, Japan, 38, 1 - 7, Mar. 2005
  • Mass production process of Neochrysocharis Formosa as the biological control against Liriomyza trifolii, 39, 41 - 54, Mar. 2006
  • Biodiversity of insect pollinators in the Nara Campus of Kinki University, 41, 77 - 94, Mar. 2008

Books etc

  • Butterfly's learning in visiting flowers-How clever are the butterflies? -,   2000

Conference Activities & Talks

  • Battus philenor larvae use their frontal projections as host plant detectors, Kandori I, Papaj DR, XXIV International Congress of Entomology,   2012 08


  • Butterfly learning during foraging for flowers, KANDORI Ikuo, Green Age, 535, 12, 15,   2018 07 , 招待有り
  • Flower learning of butterflies, especially the white cabbage butterfly, Pieris rapae, KANDORI Ikuo, Kontyu to Shizen, 51, 12, 15, 18,   2016 12 , 招待有り
  • Nectaring behaviour in the cabbage butterfly, ┣DBPieris(/)-┫DB ┣DBrapae(/)-┫DB [1] Learning abilities in visiting flowers., The Insectarium, 35, 2月, 16, 22,   1998
  • Nectaring behaviour in the cabbage butterfly, ┣DBPieris(/)-┫DB ┣DBrapae(/)-┫DB [2] Behavioural response to artificial nectar guides and its pollination effect., The Insectarium, 35, 3月, 16, 22,   1998
  • Butterfly's learning in visiting flowers., Konchu to Shizen, 34, 1, 4, 8,   1999

Research Grants & Projects

  • Ministry of Education, Culture, Sports, Science and Technology, Grants-in-Aid for Scientific Research(基盤研究(C)), Mutimodal learning in butterflies, Ikuo KANDORI, This study examined whether butterflies can learn two modalities, i.e., color and odor, during foraging for flowers. The first experiment revealed that Peris rapae butterflies can learn floral odor by dual conditioning of reward and punishment. The second experiment revealed that they also can learn floral odor by reward conditioning alone. The third experiment revealed not only that butterflies can learn floral color and odor simultaneously, but also that they can enhance the abilities to discriminate flowers by learning two modalities rather than one modality.
  • Comparison among flower visiting insects by their pollination ability and their importance as pollinators
  • Learning ability of butterflies in visiting flowers
  • Pollination ecology of dandelion, aiming for conservation of native species